Other varieties of blood oranges

Proteoglycans are proteins[1] that are heavily glycosylated. The basic proteoglycan unit consists of a "core protein" with one or more covalently attached glycosaminoglycan (GAG) chain(s).[2] The point of attachment is a Ser residue to which the glycosaminoglycan is joined through a tetrasaccharide bridge (For example: chondroitin sulfate-GlcA-Gal-Gal-Xyl-PROTEIN). The Ser residue is generally in the sequence -Ser-Gly-X-Gly- (where X can be any amino acid residue), although not every protein with this sequence has an attached glycosaminoglycan. The chains are long, linear carbohydrate polymers that are negatively charged under physiological conditions, due to the occurrence of sulfate and uronic acid groups. Proteoglycans occur in the connective tissue.

Early history (3200 BC – 625 BC)
Main articles: Tepe Sialk, Jiroft culture, Elam, and Mannaeans


19th century reconstruction of a map of the world by Eratosthenes, c.200 BC. The name Ariana (Aryânâ) was used to describe the region where the Iranian plateau is found.
Elam was part of the early urbanization during the Chalcolithic. The emergence of written records from around 3000 BC also parallels Mesopotamian history. In the Old Elamite period (Middle Bronze Age), from around 2800 BC, Elam consisted of kingdoms on the Iranian plateau, centered in Anshan, and from the mid-2nd millennium BC, it was centered in Susa in the Khuzestan lowlands. Elamite kingdom continued its existence until the emergence of Median and Achaemenid Empires.
Proto-Iranians first emerged following the separation of Indo-Iranians, and are traced to the Andronovo culture.[63] Proto-Iranian tribes arrived in the Iranian plateau in the third and second millennium BC, probably in more than one wave of emigration, and settled as nomads.
Further separation of Proto-Iranians into "Eastern" and "Western" groups occurred due to migration. By the first millennium BC, Medes, Persians, Bactrians and Parthians populated the western part, while Cimmerians, Sarmatians and Alans populated the steppes north of the Black Sea.
Other tribes began to settle on the eastern edge, as far as on the mountainous frontier of the north-western Indian subcontinent and into the area which is now Balochistan. Others, such as the Scythian tribes, spread as far west as the Balkans and as far east as Xinjiang. Avestan is an eastern Old Iranian language that was used to compose the sacred hymns and canon of the Zoroastrian Gathas in c. 1000 BC.
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